Cinder and her allies are joined in the plot to overthrow Queen Levana by the queen's stepdaughter, Winter, who has long been defying her evil stepmother's wishes by harboring feelings for her personal guard whom she is forbidden from marrying.

Vol. Two continues the team's adventures as they traverse the outer reaches of the cosmos. The Guardians must fight to keep their newfound family together as they unravel the mystery of Peter Quill's true parentage. Old foes become new allies and fan-favorite characters from the classic comics will come to our heroes' aid as the Marvel Cinematic Universe continues to expand.